Dung beetles are secondary seed dispersers. incidentally moving many of the seeds defecated by mammals vertically (seed burial) and/or horizontally as they process and relocate dung. Although several studies have quantified this ecological function of dung beetles. very few have followed seed fate until seedling establishment. https://www.rivervalleydanceac.com/product-category/t%e3%82%b7%e3%83%a3%e3%83%84/